They inhabit the Black Sea littoral from Varna to the Bosporus, the shores of the Sea of Marmora and the Aegean, the Aegean archipelago, the mainland of Greece, Epirus and the western islands as far north as Corfu.
Water of less salinity flows outwards from the Black Sea as an upper current, and water of greater salinity from the Sea of Marmora flows into the Black Sea as an under-current.
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