The Holosomata and Rhynchostomi are probably offshoots from the stem of the Araneae, and it is not unlikely (in view of the structure of the prosomatic somites of the Tartarides) that the Solifugae are connected in origin with the Pedipalpi.
The appendages of the 2nd pair were slender and pediform; those of the 3rd, 4th, 5th and 6th pairs were similar in form and ambulatory in function with their basal segments arranged round a sternal area as in the order Araneae.
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